Tessa takes up a post as a curator, Duncan takes up the role of a kept man and Richie that of a fish out of water. Duncan does the shopping, but as he is returning to the barge one morning, he sees a clown dressed as a mime kill an elderly man "For Evil's Sake".
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Helping the police, he tells them that he can identify the man. The police inspector, LeBrun, is convinced he has seen Duncan before. Later, mysterious men with guns stake out the barge and, when Duncan fells one, he finds that he is a policeman. LeBrun has remembered Duncan — when he was a young gendarme, they were chasing the assassin Kuyler and he saw Duncan then. LeBrun's assistant is in Kuyler's pay, but Duncan silences the mime forever and LeBrun arrests his assistant.
Convinced MacLeod is the gas poisoner, LeBrun tries to run him in whilst Duncan has to stop Xavier from gassing an exhibition Tessa is organising. Frustrated, Xavier comes for Duncan, but Duncan cuts off Xavier's hand and he disappears into the Seine. Grace Chandel, on the run both from the police, who want to arrest her for killing her husband, and the Immortal ex-lover who was the real killer, the possessive Carlo Sendaro "Saving Grace" was welcomed by Tessa, when Duncan and Darius help her escape, Duncan making sure that Carlo will not trouble her again.
The same was not true of Amanda, described by Duncan as "a bad habit".
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In the end, she takes Zachary's, but not before Duncan has done all the hard work. Which pretty much set the pattern for all of the Amanda stories to come. In the season finale, Duncan's old friend, Hugh Fitzcairn, comes to see him, with news that Immortals are disappearing. When they go to see Darius, they find him decapitated inside the church. Returning to Fitz's hotel, they are attacked and Fitz is spirited away, whilst Duncan returns to the church, finding a book hidden by Darius. It is a history of Immortals written by a secret society of mortals. When these mortals come for Duncan, he is ready and follows them back to their hideout, rescuing Fitz, who is about to be guillotined and confronting their leader, a wild-eyed fanatic who wants to destroy all Immortals.
He gets away, leaving a shattered MacLeod to mourn his friend and teacher.
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The series ends with MacLeod vowing revenge on the men and pouring Darius' ashes into the Seine, to flow out to the sea. This season opened with a new title sequence and a mysterious voice over explaining who Duncan MacLeod is and giving a quick summary of the Rules of the Game.
Not so mysterious a voice, as we were to find out, as it is the voice of Joe Dawson. In the first story, "The Watchers" , a bitter MacLeod returns to Seacouver, determined to find Darius' killers, but with only the Chronicle left by Darius to guide him. He follows a clue to a bookstore, where the owner, Dawson, denies knowing about the book.
When MacLeod chases down two men watching from across the street, Dawson follows and tells Duncan that he is his Watcher, one of a secret society of mortals who observe and record the history of the Immortals, but never interfere. He refuses to believe MacLeod's assertion that Darius was murdered by mortals, but is forced to when the leader of the killers is revealed to be his own brother-in-law, James Horton.
MacLeod confronts Horton, who shoots him, but not before MacLeod extracts his vengeance for Darius and runs Horton through with his sword. Believing Horton dead, but now aware of the Watchers, MacLeod carries on with his life and tries to help two old friends, but with differing results. Gregor Powers has become a photographer but has lost touch with his humanity, pushing his feelings for others away to avoid the recurring pain of loss, "Studies In Light". Duncan helps him to learn to value his life and the gift of his Immortality, whilst watching his own former lover, Linda Plager, who is mortal, die of old age and infirmity.
Soon after, another Immortal friend, Michael Moore, appears, looking for Mac's help in hunting down Quenten Barnes, Michael's long time enemy, who has reappeared after a long absence, "Turnabout". Also reappearing is Joe Dawson, who tries to enlist Duncan's help, as Barnes is hunting down and killing the men who tried and executed him years beforeTurnabout. Barnes was buried and has only recently been disinterred. An uneasy peace is struck between Watcher and Immortal, as the terrible truth dawns — Michael is Quenten Barnes, the two sides of his personality fragmented into a schizophrenic existence.
As Quenten slowly takes permanent control, Michael pleads with Duncan to end it and MacLeod kills his friend. During this episode, Mac takes Richie to a martial arts gym, or "dojo", run by Charlie DeSalvo, where they intend to train. After a spooky encounter with a palm reader reminds Mac of the gypsies' warning that he will never marry, he tempts fate by asking Tessa to marry him.
Overjoyed, she accepts and, as they plan the wedding, Mac is lured from the shop by a group of Watchers, whose leader, Pallin Wolf, stuns Richie and kidnaps Tessa. Using her as a lure, he entices MacLeod to his home, where he has a lightless room in which he hunts and kills Immortals, using a nightscope to see his prey, a la "Silence Of The Lambs". Fate smiles on Mac by allowing him to defeat Wolf and rescue Tessa despite "The Darkness", but he makes a decision that will haunt him.
Sending Tessa with Richie, he stays to rifle the man's files, but as they leave, they are mugged and shot. MacLeod runs out to find them dead in the street. As he holds Tessa in his arms, Richie revives, a newborn Immortal. Grief stricken, MacLeod sells the store. Richie arrives at the loft one night with a girl he has rescued, who was being pursued by an Immortal, "Under Color Of Authority".
From the description, Duncan recognises that the hunter is Mako, an eternal lawman. The girl is evasive, but it transpires that she is on the run for murdering her husband, who abused her, but she fears that she will be killed if she goes back, as her husband's father runs the town. Mako doesn't care — he has a warrant and will take her back to stand trial. Richie refuses to stand by and helps her to flee, but when Mako accidentally kills the girl, Richie is enraged and challenges Mako, whom he kills, despite Duncan pleading with him not to.
For the first time, Richie feels the power of a Quickening, but it is tinged with sadness as Mac tells him that he is now on his own and must leave. Duncan takes Anne to a recital by a choir of monks, one of whom, Paul, is an Immortal. Neither man realises that the song being sung is "The Song Of The Executioner", as the unseen impresario who organised the concert is Kalas, an Immortal with scores to settle with both Duncan and Paul.
Kalas kills Paul and then sets about destroying MacLeod, first by framing Joe for dealing drugs then by killing two of Anne's patients and fabricating hospital records to make it look like she is responsible. Duncan realises that Kalas is behind this and, having cleared Joe and Anne, confronts his enemy. As they fight, Anne sees Duncan fall to his death, but Richie gets her away before Duncan revives.
In the midst of this, Richie embarks on a new career in motorcycle racing. Rather than tell Anne the truth, especially with Kalas at large, Duncan leaves Seacouver, arranging for Maurice to buy back the barge in Paris. The normally philandering Fitz is in love but has a rival, who tries to kill him by running the two Immortals off the road. At the same time, Richie arrives in France, looking to build on his American successes in motorcycling. On the run, Fitz is confronted by Kalas, who beheads him as MacLeod helplessly watches.
When Duncan comes for Kalas, his nemesis escapes into a cloud of steam. Determined to finish Kalas, Duncan seeks Joe's help. Kalas owns a nightclub and Duncan "persuades" the bartender to tell him where Kalas has gone. Kalas has captured his own Watcher and is looking for information on "Methos", the legendary oldest Immortal. Duncan is stunned to find that Pierson is Methos. Kalas confronts Methos, who escapes and sets up Duncan to take his head, realising that he cannot handle Kalas, but Duncan could.
Duncan refuses, but when he confronts Kalas at Methos' apartment, their fight is interrupted by the police, who arrest Kalas for Salzer's murder after Adam Pierson identifies Kalas. Before he vanishes from Paris, Methos tells Duncan that he couldn't take the chance on Kalas winning. In the two part season Finale, Amanda arrives at the barge, being very, very nice to Duncan, which always arouses his suspicions, "Finale". Eventually, she tells him that she broke Kalas out of jail, to kill him, but that Kalas got away.
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Kalas, using the thugs employed by his cellmate, wastes no time in coming for MacLeod and, as Duncan and Amanda try to find him, finds Maurice, who Duncan rescues in the nick of time and sends out of Paris. Kalas captures Amanda, intending to use her as bait, but she escapes. A new problem arises, when Joe Dawson and Methos arrive at the barge.
Christine Salzer, the widow of Donald Salzer, the historian Kalas murdered, has found a database created for the Watchers and intends to give it to the newspapers. Dawson tries to stop her, even trying to shoot her, but MacLeod prevents him from becoming a murderer, taking the bullet himself. As the Immortals think that their secret will be on CNN, Kalas intercepts Christine and kills her, taking the vital disk. He uses the information to find the European head of the Watchers and kills him. He gives Duncan a choice — surrender to him or Kalas will broadcast the information on the disk.
Watchers and Immortals combine to hunt Kalas down. Methos tells Duncan to fight Kalas. Amanda and Duncan, faced with the destruction of their world, finally admit their feelings for each other openly. Duncan faces Kalas on the top of the Eiffel Tower and takes his enemy's head.
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The Quickening of such a powerful Immortal surges through the steelwork of the tower and overloads the Paris power grid, plunging the city into darkness and, in the process, blowing up Kalas' computer and destroying the disk. After celebrating Duncan's victory over Kalas and remembering Paul and Fitzcairn, Joe and Methos leave Duncan and Amanda alone to work out how much of what they said to each other they really want to take back.
With Kalas dead, MacLeod is free to return to his normal life and resumes his interest in antiques, until one night he sees a bracelet in a private showing at a dealer's, a bracelet he thought long buried. Shadowed by a curious Joe Dawson, he returns to his native Scotland for the first time in two hundred years to return it to the grave of his first love, Debra Campbell. Once in Glenfinnan, he is greeted with hostility by Rachel MacLeod, the local innkeeper, as the village is beset once again by mysterious murders.
With the villagers recounting the legend of Duncan MacLeod, who rose from the dead to claim his father's sword and kill the Viking raider, Kanwulf, Rachel slowly comes to trust Duncan as the legend plays itself out again and, this time, Duncan make sure he takes the Viking's head and Quickening before leaving Glenfinnan at peace.